Contribution to the knowledge of the mosses of Megriss Mountain (Algeria)
Keywords:
biodiversity; Mediterranean bryophytes; Orthotrichum scanicum; Setifian High Plains
Abstract
Due to the limited knowledge on bryophytes in Algeria, this study aimed to update the Algerian bryophyte flora. Megriss Mountain (also called Jbel or Djebel Megriss), which is a part of the High Plains of Setif, was chosen as a study area for its interesting biological and landscape diversity. The inventory was carried out in different habitats (rocks, trees, soils and streams). Samples were collected from minimum survey areas of 100 cm2 (from soil and rocks), but the sample area was sometimes increased depending on the availability of species; the trees were mostly sampled between heights of 1 and 2 meters. As a result, a preliminary list of 55 moss species were identified, including 44 acrocarpous and 11 pleurocarpous, belonging to 13 families and 29 genera. The most species-rich families were Pottiaceae, Orthotrichaceae, Brachytheciaceae, and Bryaceae, while the most diverse genera were Lewinskya and Syntrichia. Epilithic mosses were dominant, followed by terricolous mosses and finally epiphytic mosses. The most frequent species in the study area were Orthotrichum diaphanum, Lewinskya acuminate, Didymodon insulanus, Grimmia pulvinata, and Tortella squarrosa. The study also highlights the presence of a new species that had never been recorded in Algeria -Orthotrichum scanicum. A comprehensive description, microphotographs of the species are provided and its ecology is also discussed. This discovery will contribute to the enrichment of the Algerian bryophyte flora in general and of the Orthotrichaceae in particular,within the genus Orthotrichum, which now has a total of 11 species recorded in the country.References
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Bischler, H. (2004). Liverworts of the Mediterranean, ecology, diversity and distribution. Bryophytorum Bibliotheca, 61, 1–252.
Blockeel, T. L. (2012). Orthotrichum scanicum grönvall in derbyshire: An epiphytic moss new to britain. Journal of Bryology, 34(1), 55–59.
Boukhatem, A., Benslama, M., & Abed-Belouahem, D. (2017). Biodiversity and ecology of mosses in Northeastern Algeria: Case of the Watershed Tonga. Synthèse: Revue des Sciences et de la Technologie, 35, 59–68.
Boulaacheb, N. (2009). Etude de la végétation terrestre et aquatique du djebel megriss (Nord Tellien, Algérie) analyse floristique, phytosociologique et pastorale [Study of the terrestrial and aquatic vegetation of Jbel Megriss (North Tellien, Algeria) floristic, phytosociological, and pastoral analysis]. Ferhat Abbas University, Setif (in French).
Boulaacheb, N. (2013). Les paysages végétaux du djebel mégriss (Tell Septentrional, Algérie): Diversité des écosystèmes, richesse floristique, ampleur de l'anthropisation [Plant landscapes of Jbel Megriss (Northern Tell, Algeria): Diversity of ecosystems, floristic richness, extent of anthropization]. Géographie Physique et Environnement, 7, 183–210 (in French).
Camus, F. (1906). Muscinées récoltées en algérie pendant la session de la société botanique de France et déterminées [Mosses harvested in Algeria during the session of the botanical society of France and determined]. Bulletin de la Société Botanique de France, 53, 216–218 (in French).
Cano, M. J., Ros, R. M., Gallego, M. T., Jiménez, J. A., & Guerra, J. (2002). Contribution to the bryophyte flora of Morocco: The Anti-Atlas catalogue. Cryptogamie, Bryologie, 23(3), 249–262.
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Draper, I., Garilleti, R., Calleja, J. A., Flagmeier, M., Mazimpaka, V., Vigalondo, B., & Lara, F. (2021). Insights into the evolutionary history of the subfamily Orthotrichoideae (Orthotrichaceae, Bryophyta): New and former supra-specific taxa so far obscured by prevailing homoplasy. Frontiers in Plant Science, 12, 1–18.
Draper, I., Lara, F., Albertos, B., Garilleti, R., & Mazimpaka, V. (2003). The epiphytic bryoflora of the Jbel Bouhalla (Rif, Morocco), including a new variety of moss, Orthotrichum speciosum var. brevisetum. Journal of Bryology, 25, 271–280.
Draper, I., Lara, F., Albertos, B., Garilleti, R., & Mazimpaka, V. (2006). Epiphytic bryoflora of the atlas and Anti Atlas Mountains, including a synthesis of the distribution of epiphytic bryophytes in Morocco. Journal of Bryology, 28(4), 312–330.
Draper, I., Mazimpaka, V., Albertos, B., Garilleti, R., & Lara, F. (2005). A survey of the epiphytic bryophyte flora of the Rif and Tazzeka mountains (Northern Morocco). Journal of Bryology, 27, 23–34.
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Fadel, I., Magri, N., Zidane, L., Benharbit, O., Douira, A., Belahbib, N., & Dahmani, J. (2020). Contribution to the study of the bryological diversity of the Benslimane region, Morocco. Plant Archives, 20(1), 1315–1325.
Govindapyari, H., Kumari, P., Bahuguna, Y. M., & Uniyal, P. L. (2012). Evaluation of species richness of acrocarpous mosses in Imphal district, Manipur, India. Taiwania, 57(1), 14–26.
Gradstein, S. R., Churchill, S. P., & Salazar-Allen, N. (2001). Guide to the bryophytes of Tropical America. The New York Botanical Garden Press, New York.
Guerra, J., Brugués, M., Cano, M. J., & Cros, R. M. (2010). Flora briofítica ibérica Vol. 4. Sociedad Española de Briología, Murcia.
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Hodgetts, N., Söderström, L., Blockeel, T. L., Caspari, S., Ignatov, M. S., Konstantinova, N. A., Lockhart, N., Papp, B., Schröck, C., Sim-Sim, M., Bell, D., Bell, N. E., Blom, H. H., Bruggeman-Nannenga, M. A., Brugués, M., Enroth, J., Flatberg, K. I., Garilleti, R., Hedenäs, L., Holyoak, D. T., Hugonnot, V., Kariyawasam, I., Köckinger, H., Kučera, J., Lara, F., & Porley, R. D. (2020). An annotated checklist of bryophytes of Europe, Macaronesia and Cyprus. Journal of Bryology, 42(1), 1–116.
Hugonnot, V., & Celle, J. (2013). Les bryophytes du corridor alluvial de la vallée du rhône flore, végétation et fonctionnalité [Bryophytes of the Rhone alluvial complexes: Flora, vegetation and functionality]. Revue d'Ecologie (La Terre et la Vie), 68(1), 3–23 (in French).
Jelenc, F. (1955). Muscinées de l'afrique du nord: Algérie, Tunisie, Maroc, Sahara [Mosses of North Africa: Algeria, Tunisia, Morocco, Sahara]. Bulletin de la Société de Géographie et d'Archéologie de la Province d'Oran, 72–74, 1–152 (in French).
Jelenc, F. (1967). Muscinées de l’afrique du nord (supplement) [Mosses of North Africa (supplement)]. Revue bryologique et lichénologique, 35(1–4), 186–215 (in French).
Lahmar, R., Batouche, S., Labiod, H., & Meslem, M. (1993). Les sols et leur répartition dans les hautes plaines sétifiennes [Soils and their distribution in the Setif High Plains]. Eaux et Sols d’Algérie, 6, 60–70 (in French).
Laouzazni, H., Dahmani, J., Adoudou, A., & Belahbib, N. (2018). Contribution to the inventory of bryophytes of Jbel Sidi Ali in the Izarene massif (Ouazzane region, North of Morocco). American Journal of Innovative Research and Applied Sciences, 2018, 170–179.
Laouzazni, H., Zaza, J., Dahmani, J., & Belahbib, N. (2021). Contribution to the knowledge of the bryophytic flora of the izarene massif (Ouezzane region, North-West Morocco). Plant Archives, 21(1), 260–268.
Lara, F., & Mazimpaka, V. (2001). Diversité et originalité du genre Orthotrichum Hedw. (Orthotrichaceae) dans le Bassin Méditerranéen Occidental [Diversity and originality of the genus Orthotrichum Hedw. (Orthotrichaceae) in the Western Mediterranean Basin]. Bocconea, 13, 101–106 (in French).
Lara, F., Garilleti, R., & Mazimpaka, V. (1994). Clave y notas para la identificación de las especies epífitas del género Orthotrichum Hedw. En la Península Ibérica [Key and notes for the identification of the epiphytic species of the genus Orthotrichum Hedw. in the Iberian Peninsula]. Boletín de la Sociedad Española de Briología, 5, 1–6 (in Spanish).
Lara, F., Garilleti, R., Goffinet, B., Draper, I., Medina, R., Vigalondo, B., & Mazimpaka, V. (2016). Lewinskya, a new genus to accommodate the phaneroporous and monoicous taxa of Orthotrichum (Bryophyta, Orthotrichaceae). Cryptogamie, Bryologie, 37(4), 361–382.
Lara, F., Garilleti, R., Medina, R., & Mazimpaka, V. (2009). A new key to the genus Orthotrichum Hedw. in Europe and the Mediterranean region. Cryptogamie, Bryologie, 30(1), 129–142.
Medina, R., Lara, F., Goffinet, B., Garilleti, R., & Mazimpaka, V. (2013). Unnoticed diversity within the disjunct moss Orthotrichum tenellum s. l. validated by morphological and molecular approaches. Taxon, 62(6), 1133–1152.
Plášek, V., Sawicki, J., Ochyra, R., Szczecińska, M., & Kulik, T. (2015). New taxonomical arrangement of the traditionally conceived genera Orthotrichum and Ulota (Orthotrichaceae, Bryophyta). Acta Musei Silesiae, Scientiae Naturales, 64(2), 169– 174.
Rams, S., Werner, O., & Ros, R. M. (2014). Updated checklist of the bryophytes from the Sierra Nevada Mountains (S of Spain). Cryptogamie, Bryologie, 35, 261–311.
Ros, R. M., & Guerra, J. (1987). Catálogo de briófitos terrícolas de la Región de Murcia (SE de España) [Catalog of terrestrial bryophytes from the Region of Murcia (SE Spain)]. Candollea, 42, 577–599 (in Spanish).
Ros, R. M., Cano, M. J., & Guerra, J. (1999). Bryophyte checklist of Northern Africa. Journal of Bryology, 21(3), 207–244.
Ros, R. M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T. L., Brugués, M., Cano, M. J., Cros, R. M., Dia, M. G., Dirkse, G. M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J. M., Herrnstadt, I., Khalil, K., Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai, M. S., Rodríguez-Núñez, S., Sabovljević, M., Sérgio, C., Shabbara, H., Sim-Sim, M., & Söderström, L. (2007). Hepatics and anthocerotes of the Mediterranean, an annotated checklist. Cryptogamie, Bryologie, 28(4), 351–437.
Ros, R. M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T. L., Brugués, M., Cros, R. M., Dia, M. G., Dirkse, G. M., Draper, I., El Saadawi, W., Erdağ, A., Ganeva, A., Gabriel, R., González-Mancebo, J. M., Granger, C., Herrnstadt, I., Hugonnot, V., Khalil, K., Kürschner, H., Losada-Lima, A., Luís, L., Mifsud, S., Privitera, M., Puglisi, M., Sabovljević, M. S., Sérgio, C., Shabbara, H. M., Sim-Sim, M., Sotiaux, A., Tacchi, R., Vanderpoorten, A., & Werner, O. (2013). Mosses of the Mediterranean, an annotated checklist. Cryptogamie, Bryologie, 34(2), 99–283.
Smith, A. J. (2004). The moss flora of Britain and Ireland. 2nd ed. Cambridge University Press, New York.
Trabut, L. (1914). Enumération de quelques bryophytes remarquables trouvés en kabylie [Enumeration of some remarkable bryophytes found in Kabylie]. Bulletin de la Société Botanique de France, 61(5), 103–104 (in French).
Trabut, L. (1927). Quelques mousses désertiques [Some desert mosses]. Bulletin de la Société d’Histoire Naturelle d’Afrique du Nord, 18, 12–13 (in French).
Zander, R. H. (1993). Genera of the Pottiaceae: Mosses of harsh environments. Bulletin of the Buffalo Society of Natural Sciences, New York. Vol. 32.
Zaza, J., Dahmani, J., Laouzazni, H., & Belahbib, N. (2020). The bryophytic flora of the fir forest of Jbel Lekraa (Western Rif, Morocco) with census of new species. Plant Archives, 20(1), 793–800.
Zaza, J., Laouzazni, H., Dahmani, J., & Belahbib, N. (2021). Contribution to the inventory of bryophytes flora of Talassemtane National Park, North of Morocco. Plant Archives, 21(S1), 269–281.
Ben Osman, I., Hugonnot, V., Daoud-Bouattour, A., & Muller, S. D. (2021). New bryophytes in Tunisia (North Africa). Part 1: Pottiaceae. Nova Hedwigia, 113(1–2), 45–59.
Bischler, H. (2004). Liverworts of the Mediterranean, ecology, diversity and distribution. Bryophytorum Bibliotheca, 61, 1–252.
Blockeel, T. L. (2012). Orthotrichum scanicum grönvall in derbyshire: An epiphytic moss new to britain. Journal of Bryology, 34(1), 55–59.
Boukhatem, A., Benslama, M., & Abed-Belouahem, D. (2017). Biodiversity and ecology of mosses in Northeastern Algeria: Case of the Watershed Tonga. Synthèse: Revue des Sciences et de la Technologie, 35, 59–68.
Boulaacheb, N. (2009). Etude de la végétation terrestre et aquatique du djebel megriss (Nord Tellien, Algérie) analyse floristique, phytosociologique et pastorale [Study of the terrestrial and aquatic vegetation of Jbel Megriss (North Tellien, Algeria) floristic, phytosociological, and pastoral analysis]. Ferhat Abbas University, Setif (in French).
Boulaacheb, N. (2013). Les paysages végétaux du djebel mégriss (Tell Septentrional, Algérie): Diversité des écosystèmes, richesse floristique, ampleur de l'anthropisation [Plant landscapes of Jbel Megriss (Northern Tell, Algeria): Diversity of ecosystems, floristic richness, extent of anthropization]. Géographie Physique et Environnement, 7, 183–210 (in French).
Camus, F. (1906). Muscinées récoltées en algérie pendant la session de la société botanique de France et déterminées [Mosses harvested in Algeria during the session of the botanical society of France and determined]. Bulletin de la Société Botanique de France, 53, 216–218 (in French).
Cano, M. J., Ros, R. M., Gallego, M. T., Jiménez, J. A., & Guerra, J. (2002). Contribution to the bryophyte flora of Morocco: The Anti-Atlas catalogue. Cryptogamie, Bryologie, 23(3), 249–262.
Casas, C., Brugués, M., Cros, R. M., & Sérgio, C. (2006). Handbook of mosses of the Iberian peninsula. Institut d’Estudis Catalans, Barcelona.
Draper, I., Garilleti, R., Calleja, J. A., Flagmeier, M., Mazimpaka, V., Vigalondo, B., & Lara, F. (2021). Insights into the evolutionary history of the subfamily Orthotrichoideae (Orthotrichaceae, Bryophyta): New and former supra-specific taxa so far obscured by prevailing homoplasy. Frontiers in Plant Science, 12, 1–18.
Draper, I., Lara, F., Albertos, B., Garilleti, R., & Mazimpaka, V. (2003). The epiphytic bryoflora of the Jbel Bouhalla (Rif, Morocco), including a new variety of moss, Orthotrichum speciosum var. brevisetum. Journal of Bryology, 25, 271–280.
Draper, I., Lara, F., Albertos, B., Garilleti, R., & Mazimpaka, V. (2006). Epiphytic bryoflora of the atlas and Anti Atlas Mountains, including a synthesis of the distribution of epiphytic bryophytes in Morocco. Journal of Bryology, 28(4), 312–330.
Draper, I., Mazimpaka, V., Albertos, B., Garilleti, R., & Lara, F. (2005). A survey of the epiphytic bryophyte flora of the Rif and Tazzeka mountains (Northern Morocco). Journal of Bryology, 27, 23–34.
El Harech, M., Magri, N., Zidane, L., Benharbit, O., Douira, A., Belahbib, N., & Dahmani, J. (2020). Contribution to the study of bryophytic biodiversity of the Mamora forest (Morocco). Plant Archives, 20, 1305–1310.
Fadel, I., Magri, N., Zidane, L., Benharbit, O., Douira, A., Belahbib, N., & Dahmani, J. (2020). Contribution to the study of the bryological diversity of the Benslimane region, Morocco. Plant Archives, 20(1), 1315–1325.
Govindapyari, H., Kumari, P., Bahuguna, Y. M., & Uniyal, P. L. (2012). Evaluation of species richness of acrocarpous mosses in Imphal district, Manipur, India. Taiwania, 57(1), 14–26.
Gradstein, S. R., Churchill, S. P., & Salazar-Allen, N. (2001). Guide to the bryophytes of Tropical America. The New York Botanical Garden Press, New York.
Guerra, J., Brugués, M., Cano, M. J., & Cros, R. M. (2010). Flora briofítica ibérica Vol. 4. Sociedad Española de Briología, Murcia.
Guerra, J., Cano, M. J., & Brugués, M. (2014). Flora briofítica ibérica Vol. 5. Sociedad Española de Briología, Murcia.
Guerra, J., Cano, M. J., & Brugués, M. (2018). Flora briofítica ibérica Vol. 6. Sociedad Española de Briología, Murcia.
Guerra, J., Cano, M. J., & Ros, R. M. (2006). Flora briofítica ibérica Vol. 3. Sociedad Española de Briología, Murcia.
Hodgetts, N., Söderström, L., Blockeel, T. L., Caspari, S., Ignatov, M. S., Konstantinova, N. A., Lockhart, N., Papp, B., Schröck, C., Sim-Sim, M., Bell, D., Bell, N. E., Blom, H. H., Bruggeman-Nannenga, M. A., Brugués, M., Enroth, J., Flatberg, K. I., Garilleti, R., Hedenäs, L., Holyoak, D. T., Hugonnot, V., Kariyawasam, I., Köckinger, H., Kučera, J., Lara, F., & Porley, R. D. (2020). An annotated checklist of bryophytes of Europe, Macaronesia and Cyprus. Journal of Bryology, 42(1), 1–116.
Hugonnot, V., & Celle, J. (2013). Les bryophytes du corridor alluvial de la vallée du rhône flore, végétation et fonctionnalité [Bryophytes of the Rhone alluvial complexes: Flora, vegetation and functionality]. Revue d'Ecologie (La Terre et la Vie), 68(1), 3–23 (in French).
Jelenc, F. (1955). Muscinées de l'afrique du nord: Algérie, Tunisie, Maroc, Sahara [Mosses of North Africa: Algeria, Tunisia, Morocco, Sahara]. Bulletin de la Société de Géographie et d'Archéologie de la Province d'Oran, 72–74, 1–152 (in French).
Jelenc, F. (1967). Muscinées de l’afrique du nord (supplement) [Mosses of North Africa (supplement)]. Revue bryologique et lichénologique, 35(1–4), 186–215 (in French).
Lahmar, R., Batouche, S., Labiod, H., & Meslem, M. (1993). Les sols et leur répartition dans les hautes plaines sétifiennes [Soils and their distribution in the Setif High Plains]. Eaux et Sols d’Algérie, 6, 60–70 (in French).
Laouzazni, H., Dahmani, J., Adoudou, A., & Belahbib, N. (2018). Contribution to the inventory of bryophytes of Jbel Sidi Ali in the Izarene massif (Ouazzane region, North of Morocco). American Journal of Innovative Research and Applied Sciences, 2018, 170–179.
Laouzazni, H., Zaza, J., Dahmani, J., & Belahbib, N. (2021). Contribution to the knowledge of the bryophytic flora of the izarene massif (Ouezzane region, North-West Morocco). Plant Archives, 21(1), 260–268.
Lara, F., & Mazimpaka, V. (2001). Diversité et originalité du genre Orthotrichum Hedw. (Orthotrichaceae) dans le Bassin Méditerranéen Occidental [Diversity and originality of the genus Orthotrichum Hedw. (Orthotrichaceae) in the Western Mediterranean Basin]. Bocconea, 13, 101–106 (in French).
Lara, F., Garilleti, R., & Mazimpaka, V. (1994). Clave y notas para la identificación de las especies epífitas del género Orthotrichum Hedw. En la Península Ibérica [Key and notes for the identification of the epiphytic species of the genus Orthotrichum Hedw. in the Iberian Peninsula]. Boletín de la Sociedad Española de Briología, 5, 1–6 (in Spanish).
Lara, F., Garilleti, R., Goffinet, B., Draper, I., Medina, R., Vigalondo, B., & Mazimpaka, V. (2016). Lewinskya, a new genus to accommodate the phaneroporous and monoicous taxa of Orthotrichum (Bryophyta, Orthotrichaceae). Cryptogamie, Bryologie, 37(4), 361–382.
Lara, F., Garilleti, R., Medina, R., & Mazimpaka, V. (2009). A new key to the genus Orthotrichum Hedw. in Europe and the Mediterranean region. Cryptogamie, Bryologie, 30(1), 129–142.
Medina, R., Lara, F., Goffinet, B., Garilleti, R., & Mazimpaka, V. (2013). Unnoticed diversity within the disjunct moss Orthotrichum tenellum s. l. validated by morphological and molecular approaches. Taxon, 62(6), 1133–1152.
Plášek, V., Sawicki, J., Ochyra, R., Szczecińska, M., & Kulik, T. (2015). New taxonomical arrangement of the traditionally conceived genera Orthotrichum and Ulota (Orthotrichaceae, Bryophyta). Acta Musei Silesiae, Scientiae Naturales, 64(2), 169– 174.
Rams, S., Werner, O., & Ros, R. M. (2014). Updated checklist of the bryophytes from the Sierra Nevada Mountains (S of Spain). Cryptogamie, Bryologie, 35, 261–311.
Ros, R. M., & Guerra, J. (1987). Catálogo de briófitos terrícolas de la Región de Murcia (SE de España) [Catalog of terrestrial bryophytes from the Region of Murcia (SE Spain)]. Candollea, 42, 577–599 (in Spanish).
Ros, R. M., Cano, M. J., & Guerra, J. (1999). Bryophyte checklist of Northern Africa. Journal of Bryology, 21(3), 207–244.
Ros, R. M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T. L., Brugués, M., Cano, M. J., Cros, R. M., Dia, M. G., Dirkse, G. M., El Saadawi, W., Erdağ, A., Ganeva, A., González-Mancebo, J. M., Herrnstadt, I., Khalil, K., Kürschner, H., Lanfranco, E., Losada-Lima, A., Refai, M. S., Rodríguez-Núñez, S., Sabovljević, M., Sérgio, C., Shabbara, H., Sim-Sim, M., & Söderström, L. (2007). Hepatics and anthocerotes of the Mediterranean, an annotated checklist. Cryptogamie, Bryologie, 28(4), 351–437.
Ros, R. M., Mazimpaka, V., Abou-Salama, U., Aleffi, M., Blockeel, T. L., Brugués, M., Cros, R. M., Dia, M. G., Dirkse, G. M., Draper, I., El Saadawi, W., Erdağ, A., Ganeva, A., Gabriel, R., González-Mancebo, J. M., Granger, C., Herrnstadt, I., Hugonnot, V., Khalil, K., Kürschner, H., Losada-Lima, A., Luís, L., Mifsud, S., Privitera, M., Puglisi, M., Sabovljević, M. S., Sérgio, C., Shabbara, H. M., Sim-Sim, M., Sotiaux, A., Tacchi, R., Vanderpoorten, A., & Werner, O. (2013). Mosses of the Mediterranean, an annotated checklist. Cryptogamie, Bryologie, 34(2), 99–283.
Smith, A. J. (2004). The moss flora of Britain and Ireland. 2nd ed. Cambridge University Press, New York.
Trabut, L. (1914). Enumération de quelques bryophytes remarquables trouvés en kabylie [Enumeration of some remarkable bryophytes found in Kabylie]. Bulletin de la Société Botanique de France, 61(5), 103–104 (in French).
Trabut, L. (1927). Quelques mousses désertiques [Some desert mosses]. Bulletin de la Société d’Histoire Naturelle d’Afrique du Nord, 18, 12–13 (in French).
Zander, R. H. (1993). Genera of the Pottiaceae: Mosses of harsh environments. Bulletin of the Buffalo Society of Natural Sciences, New York. Vol. 32.
Zaza, J., Dahmani, J., Laouzazni, H., & Belahbib, N. (2020). The bryophytic flora of the fir forest of Jbel Lekraa (Western Rif, Morocco) with census of new species. Plant Archives, 20(1), 793–800.
Zaza, J., Laouzazni, H., Dahmani, J., & Belahbib, N. (2021). Contribution to the inventory of bryophytes flora of Talassemtane National Park, North of Morocco. Plant Archives, 21(S1), 269–281.
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